For each tower up he deals 25% more damage. It's a cumulative buff rather than a linear one: so for 1 tower up he'll do 125% of his normal damage, for 2 towers up he'll do 125%+(25% of 125%), so with 3 towers up he'll deal 196% of his damage, but with 4 towers up 245% and when he pops his cooldown (forgot the name of the spell, where he does increased dmg), that becomes pretty nuts, definitely when you have the adds dealing cleave damage as well if the raid doesn't move out.
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