With both Rune Strike and Maul the abilties used to affect the next melee swing. I don't think that this is still the case. When it affected the next swing the ability was a toggle and the ! was necessary to prevent the ability from being toggled on and then off if you hit the macro twice before your next melee swing.
I am fairly certain that this is no longer the case with Rune Strike, not sure about Maul.
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